THE BOTTOM FAUNA OF THE BLACKWATER ESTUARY 19 Fig. 3. The substrates of the study area on an aerobic mud base, with a considerable quantity of similar shell-remains mixed with the mud. The stations due north of the wall possessed, in general, more and cleaner shells than those to the north-east and north-west, which were characterised by a gritty shell and mud mix with little surface whole-shell. The remaining stations, to the south of the wall, had a sub- strate composed of a layer of soft aerobic mud (of up to 10cm thickness) overlying hard black mud (of up to 20cm thickness) and stiff clay (which was reached from 0.5 to 25cm below the surface). The western area was often floored by almost bare grey clay, but towards the east the layers of mud and silt increased in thickness. Surface shell was absent. THE NATURE OF THE FAUNA Since samples were sorted by hosing over a 3mm mesh sieve, small or fragile species were not retained and all such species were discounted with respect to the present surveys. In particular in this context, amphipods were seldom retained and were ignored. In total, fifty-one species (including the "colonial hydroid" category as a single species) were recorded and a further two "incidental" species (the small or fragile species re- ferred to above) were identified. These species are listed in Table 1. They include ten species not recorded by the general survey of 1970, of which three are not listed by Davis (1967) and are therefore new records for the Blackwater (these are asterisked in that table, and see also Appendix 1). In addition,