36 THE ESSEX NATURALIST Results and discussion of methods Table 4 summarises the relative abundance of the small mammals in the main surveys described above. The fact that one species (the Dormouse, Muscardinus avellanarius) was not detected by any of these means is commented on further under that species heading below. Table 4 A comparison of the results from four small mammal sampling methods a. The extensive live-trapping survey (see appendix 1) b. The intensive live-trapping survey at Coptfold (see table 1) c. The bottle hunts (see table 2) d. Owl pellets (see table 3) The great differences between the results from the three main methods, and the fact that each method revealed a different species as the 'most abundant' indicates the degree of caution necessary when interpreting the results of such surveys. Clearly each method gives a biased catch and some of the reasons for this are as follows — Live-trapping. To avoid theft of traps, woodlands or hedgerows were usually selected as trap-sites. Bottles and Owls sample different habitats. Also the overnight trapping and the absence of pre-baiting is likely to increase the proportion of Apodemus spp. Voles tend to be trap shy and both shrews and voles are partially diurnal. The very lightest species are likely to be underestimated by trapping as a certain pressure is necessary to fire the mechanism. Species such as Harvest Mice (Micromys minutus) which live above ground level for part of the year will be underestimated for this reason. Bottles. Most bottles are dumped in roadside hedgerows and lay-bys. Very few were collected from mature woodland. Since the bottles trap over a long period they may detect species which migrate through the area—these could be missed by short-term live-trapping. The bottles are not biased against light species and the narrowest necked bottles would only allow the smaller species and individuals to enter. Studies by Margaret Wise at Royal Holloway College have shown that different