12 A shrub/tree layer occurred in the marsh area and included essentially young trees, e.g. willow (Salix sp.), ash (Fraxinus excel- sior} , beech (F. sylvatica) and alder (Alnus glutinosa) , with large mature trees on the margins of the area (including oak (Q. robur), hawthorn (Crataegus monogyna), birch (B. pendula) , and beech (F. sylvatica) . Coarse tussocks of vegetation occurred over the area and included such species as cotton grass (Eriophorum sp.), spike rush (Eleocbaris palustris), willowherbs (Epilobium hirsutum, E. parviflorum), marsh bedstraw (Galium palustre), marsh thistle (Cirsium palustre), sharp flowered rush (Juncus acutiflorus) , as well as ragwort (Senecio jacobaea) on the drier marginal areas. Many of these plants are indicative of a poorly drained acidic soil, and this supports the observations made in the soil pro-files of the site. 3.2. Habitat Factors For a general study of this kind, it was not practicable to try to analyse all the climatic, edaphic and biotic factors. It was considered more important to generally assess those -factors which appear to play a prominent part in the -factor complex. Temperature The air temperature was recorded at each site approximately 5 feet above ground level (together with the water temperature at site 3) at between 10.45 and 12.30 hours at each visit (Table 1.). Ideally a greater number of temperature readings should have been taken at