from areas beyond the limits of the Forest sensu Buck, include at least 38 ad-
ditional to those listed by Buck (1955) or in Table 5. A number of these are very
early records for "Epping" and its district by Doubleday (1836):- Rantus
aberratus Gemm. & Har., R. exsoletus (Forster), Dytiscus semisulcatus Muller,
Berosus affinis Brulle, B. luridus (Linn.), Oiceoptoma thoracicum (Linn.),
Trox sabulosus (Linn.), Ampedus cardinalis (Schiodte), Strangalia melanura
(Linn.), Callidium violaceum (Linn.) and Hylotrupes bajulus (Linn.). Species
taken by E. A. Newbery at Highams Park, but unknown from Epping Forest
sensu Buck are:- Clivina collaris (Herbst), Trechus micros (Herbst), Atheta
celata (Er.), Aloconota insecta (Thoms.) and Psylliodes napi (Fabr.). Four
species taken at Walthamstow towards the end of the nineteenth century, in
association with wood ants, by Butler, have not yet been found in the main body
of the Forest. These are Myrmetes piceus (Payk.), Dendrophilus pygmaeus
(Linn.), Monotoma conicicollis Aube and M. angusticollis Gyll., nineteenth
century records for further species not known from the Forest sensu Buck in-
clude Teretrius fabricii Mazur, Stenus nanus Steph., Xantholinus angularis
Ganglb., Velleius dilatatus (Fabr.), Neuraphes sparshalli (Denny), Oligota
flavicornis (Boisd. & Lac), Bruchidius ater (Marsh.), Ampedus cardinalis
(Schirodte) and Aphodius porcus (Fabr.), all at Wanstead, and Leptacinus
fornicetorum Markel at Wood St., Epping. Incidentally, records of
Onthophilus punctatus (Muller) and Agrilus viridis (Linn.) (see Levey, 1977) for
"Woodford" (December 1864, J. Power) are quite likely to relate to specimens
taken at some other locality, as Power's diaries (in BM(NH)) make it clear that
he was given the specimens, by Desvignes, when on a visit to Woodford. 20th
century records for areas not covered by Buck (1955) include Lesteva pubescens
Mann. at Walthamstow, Quedius fulgidus (Fabr.) and Cilea silphoides (Linn.)
at Woodford, Brachypterolus vestitus (Kies.) at Wanstead, Helops caeruleus
(Linn.) from "near Epping", Metoecus paradoxus (Linn.) from "Chingford",
Bembidion quadripustulatum Serv. and Graphoderus cinereus (Linn.) at
Upshire. All of these are unknown from the Forest sensu Buck. The identity of
Upshire specimens of the last-named species has been checked, and they prove
to be the true Graphoderus cinereus, rather than either of the two other species
of the genus now known to be British.
Other areas which historically formed part of the Forest of Essex, but are
not included in the Epping Forest of today are also known to support interesting
beetle faunas. For example, some twenty beetle species unknown from the
Forest proper are known from the woodlands (Coopersale Common, etc.) which
adjoin the Lower Forest, although several of these are associated with planted
conifers found in Ongar Park Wood.
DISCUSSION OF THE FOREST BEETLE FAUNA
The beetle species known from Epping Forest sensu Buck (1955) thus
number in the region of 1332 (a few of these resting on rather slender evidence),
while a further 39 species may be added to this list if other areas of Epping
48