mixta (the Migrant Hawker) and Orthetrum cancellatum appear to be expanding in range and abundance from strongholds in the south-east. Dragonfly Behaviour On emergence from the aquatic habitats in which they spend the early stages of their life-histories, the adult dragonflies make a 'maiden' flight directly away from the emergence site. Some subsequently return to the site of origin, but more commonly there is a general dispersal of individuals. The aquatic habitats of dragonflies are in any case often transitory because of natural ecological succession, or because of human intervention. The ability to disperse, locate and colonise new breeding sites is therefore essential to the long-term survival of most species. After emergence some two to three weeks may be spent away from any suitable breeding site. During this period some species, such as the large hawkers of genus Aeshna may be seen hawking or basking in woodland rides or glades, well away from water. The less powerful damselflies are generally to be found closer to their breeding sites. When a suitable breeding site has been located many species show some territorial activity. The males of several hawkers (such as Anax imperator, the Emperor Dragonfly), darters (such as Sympetrum sanguineum, the Ruddy Darter), and damselflies (such as Enallagma cyathigerum, the Common Blue Damselfly) may show quite aggressive territoriality. Presumably such behaviour maximises mating opportunities for males which successfully hold territories, and also secures 'choice', egg-laying sites for females. Since it limits the number of breeding individuals in any one site, it presumably also results in more effective dispersal of the population to other sites. Only in Calopteryx species among the British dragonflies is there any elaborate 'courtship' activity, but all species share the distinctive 'wheel' mating posture (Fig. 2:3). Fig 2.3 'Wheel' Mating Posture (in Enallagma cyathigerum) 15