THE ESSEX NATURALIST Journal of the ESSEX FIELD CLUB No 16 (New series) For the year 1998/99 ISSN 0071-1489 Editorial team: Dr Ken Adams, Dr Peter Allen, Dr Jeremy Dagley, Dr Chris Gibson, Mr Peter Harvey (Technical Editor) Copyright © Essex Field Club 1999 All rights reserved. No part of this publication may be reproduced, stored in a retrieval system or transmitted, in any form or by any means, electronic, mechanical, photocopying, recording or otherwise. without the permission of the Essex Field Club or the copyright owner. Information published in this journal may be freely quoted provided that full acknowledgement is made in the normal manner to both author and journal. Published by the Essex Field Club c/o Department of Life Sciences, University of East London, Romford Road, Stratford, London E15 4LZ Printed by The Book Company, Ipswich