Utilisation of Molinia caerulea Tussocks by Overwintering Beetles at Epping Forest 117 Assuming that the cores were representative of whole tussocks, it can be concluded that some aspect of the staphylinids allowed them to successfully utilise the flooded tussocks, and /or that some aspect of the carabids excluded them. Two - Way Analysis of Variance revealed an overall significant difference between water content of the cores (P