Millenial Status in Essex of Nationally Scarce and Threatened Species of Coleoptera 189 Discussion The data contained in Table 1 along with data on high scarcity/threat status species dealt with in the previous paper (Hammond, 1999) are summarised below in Table 3. Excluding the 64 species now considered to be extinct in the U.K., the summarised data reveal that some 45% of the beetle species considered to merit a national scarcity /threat status have been reliably recorded, at one time or another, from Essex. 37% of the scarce/threatened British species have been reliably recorded from the county since 1900, 27% since 1949, and just under 20% since 1969. A very subjective assessment of these data suggests that had no species actually present in the county remained undetected, these figures would be substantially higher. Indeed, although only 341 of the scarce/threatened species have been reliably reported from Essex since 1969, the number of these species still present in the county is quite likely to be double that figure. To select a few scattered examples, it would be very surprising if such species as Omalium rugatum, Philonthus pseudoparcus, Oxypoda amoena, O. recondita, Meligethes incanus, Synchita humeralis, Cis festivus, Phaedon concinnus and Xyleborus dispar were not still to be found in Essex. However, the task of investigating and confirming the continuing occurrence of these species in the county is a considerable one, as many of them are extreme habitat or microhabitat specialists not likely to be detected by every-day collecting techniques. Not surprisingly, the picture painted by the summary presented in Tabic 3 differs in detail from one category of species to another. For example, some 21% of U.K. beetle species in the highest category (RDB1) have been reliably reported from Essex but only 3% or so of RDB1 species have been found here in the most recent (post-1969) period. The picture for lower status species is very different. For example, 69% of the lowest status (Nb) species have been recorded from Essex and, despite the inadequacy of the recording effort, 37% of the U.K. species categorised 'Nb' have been reliably reported in the county since 1969. Table 3. Coleoptera species with high conservation status. The Essex figures omit those species (in square brackets in Table 2 and in square brackets in Hammond, 1999: Table 1) for which records are particularly doubtful. RDB1 RDB2 RDB3 RDBI RDBK Extinct Na Nb N Total U.K.total 154 57 114 122 250 64 259 548 234 1802 Essex total 34 18 58 15 54 7 124 379 108 797 Essex 1900+ 18 14 50 12 43 - 96 330 92 655 Essex 1950+ 11 8 42 4 25 - 68 266 62 486 Essex 1970+ 5 4 22 2 21 - 49 205 33 341 Any very detailed analysis of the results presented here in terms of what they may indicate with respect to future recording efforts and their possible implications for conservation efforts is scarcely warranted by the quality of the data. Suffice it to say on the first count that many different parts of the county, habitat types and microhabitats are in need of further investigation if a really accurate picture of the current situation viz-a-viz beetle species in Essex is to be obtained. The use of specialist collecting techniques (e.g. for sampling the subterranean fauna), and of modern devices (e.g. large- area flight interception traps that are effective in sampling species living in "cryptic" situations such as the fruiting bodies of truffles and slime-moulds) will be of great service in achieving this aim. To the extent that the data summarised here do give a reasonable idea of which elements of the beetle fauna have suffered the greatest declines and may be under the greatest continuing threat, a few final Essex Naturalist (New Series) 17 (2000)